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VST System Link: VST System Link is Steinberg's platform-independent technology for using Cubase with other computers via digital audio interface connections running System Link-compatible software, such as Cubase SX/SL, Cubase VST 5.2 (which runs in slave mode only), and Nuendo. If you're using Cubase on a network of computers running VST System Link-compatible software and you want to sync Cubase to the master machine via VST System Link, this is the option to enable.
The interesting thing about VST System Link is that it also includes timing and tempo information, which is important because unlike Cubase VST, SX/L can't be slaved to MIDI Clock, due to the newer timing engine being based around linear time rather than musical time. This means that if you want to set up a Cubase system primarily as a mixer (slaved to another Cubase system, for example) with tempo-dependent effects, VST System Link is your only choice.